Chemicals ↓ Chemotrophy Phototrophy Organic Inorganic chemicals chemicals (glucose,acetate,etc.) (H2,H2S,Fe2+,NH4",etc.) Chemoorganotrophs Chemolithotrophs Phototrophs (glucose +2-CO2+H2O)(H2+02-H2O)(lightATP) ATP ATP
第三节能量来自光 产氧光合作用 不产氧光合作用 以视紫红质为基础的光合作用
第三节 能量来自光 产氧光合作用 不产氧光合作用 以视紫红质为基础的光合作用
Table 9.5 Diversity of Phototrophic Organisms Eucaryotic Organisms Procaryotic Organisms Plants Cyanobacteria Multicellular green,brown, Green sulfur bacteria and red algae Green nonsulfur bacteria Unicellular protists Halobacterium (archaeon) (e.g.,euglenoids, Purple sulfur bacteria dinoflagellates,diatoms) Purple nonsulfur bacteria Prochloron oxygenic photosynthesis-eucaryotes and cyanobacteria anoxygenic photosynthesis -all other bacteria
oxygenic photosynthesis – eucaryotes and cyanobacteria anoxygenic photosynthesis – all other bacteria Table 9.5
Chlorophyll-based phototrophy Light Chlorophyll or Electron transport chain NAD(P)+ bacteriochlorophyll NAD(P)H photo phos PMF ATP Carbon source (often CO2) Biosynthesis Rhodopsin-based phototrophy Light Bacteriorhodopsin photo phos (archaeorhodopsin) PMF ATP proteorhodopsin Carbon source 。Biosynthesis
CH H 0.9 0.8 Chlorophyll a Green alga C, peaks Chlamydomonas 0.7 Carotenoid 0 peak 衣藻 HgC 05 H 0.4 H2C H 0.3 CH2 H COOCH3 0.2 Cyclopentanone COOC20H39 ring 0.1 Phytol 0 340400 500600700 800900 Chlorophyll a Wavelength(nm) 0.9 Bacteriochlorophyll a peaks H HC H 8 Carotenoid 、H 0.7 peaks 0.6 Purple bacterium 0.5 Rhodopseudomona H C H CH3 0.3 红假单胞菌 H2C H CH2 HT d 0.2 COOCH 0.1 Cyclopentanone COOC2oH39 ring 0 Phytol 40400500600700800900 Wavelength(nm) Bacteriochlorophyll a
Green alga Chlamydomonas Purple bacterium Rhodopseudomona s 红假单胞菌 衣藻